New records of Coleoptera from Lagodekhi Protected Areas, with new records for Georgia (Sakartvelo)

George Japoshvili; Jacek Hilszczański; Adam Byk; Tomasz Jaworski; Krzysztof Łoś; Jerzy Borowski; Grzegorz Tarwacki; Jacek Piętka; Radosław Plewa

doi:10.3897/caucasiana.1.e85239

Abstract

During this survey, 195 species of coleopterans have been recorded from the Lagodekhi Protected Areas (LPA). Thirty families are recorded for the first time from this area, of which Rhysodidae, Sphindidaeand, and Thymalidae are new for Georgia. One hundred fifty-seven species are new to LPA and 66 of them are new country records.

Key words

Beetles, South Caucasus, Faunistics, Biodiversity Inventory

Introduction

The Lagodekhi Natural Reserve was established in 1912 (thanks to the efforts of the Polish forester and naturalist Ludwik Młokosiewicz (Chodubski 1982)), and is located in the extreme north-eastern part of Georgia (southern slopes of the Caucasus Mountains) with a range of altitudes from 590 to 3500 m above sea level. The Lagodekhi Protected Areas (LPA) include the Lagodekhi Natural Reserve (19,749 ha) and a Managed Reserve (4702 ha) (APA 2016). The LPA keeps one of the world's best-preserved primeval deciduous forest habitats (Fig. 1A). The first summarized paper about the insect diversity of Lagodekhi Natural Reserve was given by Kobakhidze (1956), listing 632 insect taxa (Japoshvili et al. 2022). A very recent update about the insect diversity of LPA provides 1790 species records for LPA (Mumladze et al. 2017; Japoshvili 2017, 2022; Japoshvili et al. 2022). This makes the LPA the best studied protected area (by means of insect diversity) in the Caucasus Region. However, the true diversity of insect species in LPA is still far from being fully inventoried. For instance, materials collected in 2014 revealed 313 new species records for the family Ichneumonidae (Hymenoptera) for the LPA, including 209 new country records (Riedel et al. 2018). The coleopteran insects of LPA are relatively well known thanks to the old works by a number of authors (Chantladze 2000; Jambazishvili 2000; Cholokava et al. 2002; Javelidze et al. 2002; Merkviladze and Kvavadze 2002; Reck and Chaladze 2004; Cholokava 2008). However, Coleoptera, which is one of the most specious orders of insects, is also supposedly represented by many more species in LPA than we know today. A recent work by Gul Aslan et al. (2017) who studied malaise trap material of leaf beetles (Chrisomelidae) collected in 2014, reported 48 species with 14 new country records. Here we provide inventory data for a number of different families of Coleoptera, collected within the same malaise trap program mentioned above, and supplemented by other materials collected in recent years.

Figure 1.

A – Forest habitat near the main entrance of Lagodekhi National Park; B – Malaise trap in the mountain middle zone in a mixed mountain forest, with Fagus orientalis as a dominant tree species (Fageto-galiosum), 1351 m a.s.l.; C – Funnel and yellow pan traps, close to river along the road from main entrance to Lagodekhi National Park; D – Strongly decomposed beech snag; E – Snag of beech with emergence holes of Rhaesus serricollis (Motschulsky, 1838) (Cerambycidae); F – Bracket fungus with beetles of Diaperis boleti (Linnaeus, 1758) (Tenebrionidae).

Materials and methods

The materials represented in the present article were collected by the first author using malaise traps (MT) operating in LNP during the entire growing season from April to November of 2014. However, at high altitudes (H5, H6 – see below), collecting began later (5 May 2014; alpine 23 May 2014) and was completed earlier (6 October 2014), due to climate conditions at higher altitudes. MTs (Fig. 1B) were placed at the following sites: H1: low-altitude forest, 666 m above sea level (a.s.l.) (N41.852483, E46.287767); H2: mid-altitude forest, 847 m a.s.l. (N41.854183, E46.292733); H3: high-altitude forest, 1351 m a.s.l. (N41.871467, 46.311533); H4: subalpine forest, 1841 m a.s.l. (N41.882733, 46.32185); H5: subalpine meadow and scrubland, 2230 m a.s.l. (N41.89805, E46.333883); H6: alpine zone, 2559 m a.s.l. (N41.906183, E46.3334). Before being sorted, samples were collected every 10 (± 2) days and kept in 96% ethanol. Additionally, 12-funnel traps (FT hereafter), from 1 May to 5 August 2014, and Moericke traps (yellow pan traps; hereafter - PT), from 1 May to 14 June 2014, have been used to supplement the MT collection (Fig. 1C). FTs and PTs were set up in the same localities as MTs. Finally, our material also contains data obtained using occasional hand collection (HC hereafter) during the various field research campaigns in LPA during the last decade. In such cases, geographic coordinates are given under each of the species listed below.

The following authors have helped to identify materials: S. Mazur (Histeridae), J. Borowski (Ptinidae), W.H. Rücker and R. Plewa (Latridiidae), A. Lasoń (Nitidulidae), A. Melke (Staphylinidae), R. Królik (Ciidae and Buprestidae), M. Wanat (Curculionidae (without Scolytinae)), J. Mertlik and G. Platia (Elateridae), and T. Gazurek (various taxa). The rest of the material was identified by the authors. All new records for LPAs are marked with one asterisk (*) and new country records with double asterisks (**). New country records were verified mainly based on the Catalogue of Palaearctic Coleoptera (Löbl and Smetana 2006, 2007, 2011, 2013; Löbl and Löbl 2015, 2016, 2017; Danilevsky 2020; Iwan and Löbl 2020). In cases when we were not able to find an original source for the Georgian distribution of a particular taxon with no later confirmation, we did not take into consideration such records. The voucher materials are placed in the author’s collections. The abbreviated names of the collectors are as follows: George Japoshvili – GJ; Radosław Plewa – RP; Jacek Hilszczański – JH; Jerzy Borowski – JB; Jacek Piętka – JP; Adam Byk – AB; Tomasz Jaworski – TJ; Krzysztof Łoś – KL. If the material was gathered by a group of people, we use the abbreviation IPT (IBL: Forest Research Institute (Instytut Badawczy Lenictwa) Polish Team). Material for correct terminology was checked according to the Global Biodiversity Information Facility (GBIF 2022).

Results

In the species list given below, the information is provided in the following order: the number of specimens, sampling locality, sampling date, collection method, and collectors.

Order: Coleoptera

Family: Anthribidae*

Genus: Dissoleucas Jordan, 1925

Dissoleucas niveirostris (Fabricius, 1798)

GEORGIA • 2 inds; H1; 3.05.2014; MT. 1 ind; H1; 15-25.05.2014; MT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT.

Genus: Platyrhinus Clairville, 1798

Platyrhinus resinosus (Scopoli, 1763)**

GEORGIA • 1 ind; N41.84, E46.28; 29.05.2012; HC; leg: RP.

Genus: Platystomos Schneider, 1791

Platystomos albinus (Linnaeus, 1758)

GEORGIA • 1 ind; H1; 3.05.2014; MT.

Genus: Pseudeuparius Jordan, 1914

Pseudeuparius sepicola (Fabricius, 1793)

GEORGIA • 1 ind; H3; 4.05.2014; MT. 1 ind; N41.84, E46.28; 4.05.2014; HC; leg: RP.

Genus: Choragus Kirby, 1819

Choragus sheppardi (Kirby, 1819)

GEORGIA • 2 inds; H2; 5-14.07.2014; FT; leg: IPT.

Family: Biphyllidae*

Genus: Diplocoelus Guérin-Méneville, 1838

Diplocoelus humerosus Reitter, 1876

GEORGIA • 2 inds; H1; 5-15.07.2014; MT.

Family: Buprestidae*

Genus: Anthaxia Eschscholtz, 1829

Anthaxia signaticollis Krynicki, 1832

GEORGIA • 1 ind; H2; 1.05-14.06.2014; PT; leg: IPT.

Genus: Coraebus Laporte de Castelnau & Gory, 1839

Coraebus rubi (Linnaeus, 1767)

GEORGIA • 1 ind; N41.85, E46.23; 2.07.2011; HC; leg: JH.

Genus: Dicerca Eschscholtz, 1829

Dicerca chlorostigma Mannerheim, 1837

GEORGIA • 1 ind; N41.85, E46.28; 30.05.2012; HC; leg: JH. 1 ind; H3; 25.05-4.06.2014; MT. 1 ind; H3; 15-25.06.2014; MT. 1 ind; H1; 26.07-5.08.2014; MT. 1 ind; H1; 5-15.08.2014; MT.

Genus: Chrysobothris Eschscholtz, 1829

Chrysobothris affinis (Fabricius, 1794)

GEORGIA • 1 ind; H5; 15-25.06.2014; MT.

Genus: Lamprodila Motschulsky, 1860

Lamprodila rutilans (Fabricius, 1777)

GEORGIA • 1 ind; 2.07.2011; HC; leg: JH.

Family: Carabidae

Genus: Carabus Linnaeus, 1758

Carabus septemcarinatus Motschulsky, 1840*

GEORGIA • 1 ind; N41.85, E46.28; 1.05.2014; HC; leg: KL.

C. staehlini Adams, 1817

GEORGIA • 1 ind; H1; 17-23.06.2015; HC; leg: GJ.

Genus: Tachyta Kirby, 1837

Tachyta nana (Gyllenhal, 1810)

GEORGIA • 3 inds; H1; 23.04-3.05.2014; MT. 1 ind; H1; 3.05.2014; MT.

Genus: Aegosoma Audinet-Serville, 1832

Aegosoma scabricorne (Scopoli, 1763)

GEORGIA • 1 ind; N41.84, E46.28; 30.04.2014; at night on snag; HC; leg: JH; 2 inds; N41.84, E46.28; 17-21.07.2019; HC; leg: JB, AB, JP.

Family: Cerambycidae

Genus: Agapanthia Audinet-Serville, 1835

Agapanthia lederi Ganglbauer, 1884*

GEORGIA • 1 ind; H6; 25.06-5.07.2014; MT. Genus: Alosterna tabacicolor (Degeer, 1775)

Alosterna tabacicolor subvittata Reitter, 1885*

GEORGIA • 1 ind; H3; 15-25.06.2014; MT. 1 ind; H3; 4-14.06.2014; MT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT.

Genus: Anaglyptus Mulsant, 1839

Anaglyptus simplicicornis Reitter, 1906*

GEORGIA • 2 inds; H3; 4-14.06.2014; FT. leg: IPT.

Genus: Aromia Audinet-Serville, 1834

Aromia moschata ambrosiaca Steven, 1809

GEORGIA • 7 inds; N41.84, E46.28; 17-21.07.2019; HC; leg: JB, AB, JP.

Genus: Cerambyx Linnaeus, 1758

Cerambyx cerdo acuminatus Motschulsky, 1853

GEORGIA • 4 inds; N41.84, E46.28; 17-21.07.2019; HC; leg: JB, AB, JP.

Genus: Chlorophorus Chevrolat, 1863

Chlorophorus varius (Müller, 1766)*

GEORGIA • 1 ind; N41.85, E46.23; 2.07.2011; HC; leg: JH.

Genus: Clytus Laicharting, 1784

Clytus schneideri Kiesenwetter, 1879*

GEORGIA • 3 inds; N41.85, E46.23; 2.07.2011; HC; leg: JH.

Genus: Cortodera Mulsant, 1863

Cortodera pumila Ganglbauer, 1882*

GEORGIA • 1 ind; N41.85, E46.23; 2.07.2011; leg: JH. 2 inds; H3; 4-14.06.2014; FT. 1 ind; H3; 5-15.05.2014; MT. 1 ind; H3; 15-25.05.2014; MT.

Genus: Exocentrus Dejean, 1835

Exocentrus adspersus Mulsant, 1846*

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT. 1 ind; H1; 15-25.07.2014; MT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT.

Genus: Fallacia Mulsant & Rey, 1863

Fallacia elegans (Faldermann, 1837)*

GEORGIA • 1 ind; H1; 12-23.04.2014; MT. 1 ind; H3; 15-25.05.2014; MT.

Genus: Leiopus Audinet-Serville, 1835

Leiopus femoratus Fairmaire, 1859*

GEORGIA • 2 inds; H3; 4-14.06.2014; FT; leg: IPT.

L. nebulosus caucasicus Ganglbauer, 1887*

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT. 1 ind; H3; 15-25.06.2014; MT.

Genus: Leptura Linnaeus, 1758

Leptura quadrifasciata lederi Ganglbauer, 1882*

GEORGIA • 2 inds; N41.85, E46.23; 2.07.2011; leg: JH; 1 ind; H1; 25.06-5.07.2014; MT. 4 inds; H1; 17-21.07.2019; HC; leg: JB, AB, JP.

Genus: Mesosa Latreille, 1829

Mesosa curculionoides (Linnaeus, 1760)

GEORGIA • 1 ind; H1; 15-27.09.2014; MT.

Genus: Morimus Brullé, 1832

Morimus verecundus (Faldermann, 1836)*

GEORGIA • 1 ind; N41.84, E46.28; 30.04.2014; HC; leg: JH. 1 ind; H1; 15-25.06.2014; MT. 1 ind; H1; 5-15.08.2014; MT. 2 inds; H1; 17-21.07.2019; HC; leg: JB, AB.

Genus: Nathrius Brethes, 1916

Nathrius brevipennis (Mulsant, 1839)*

GEORGIA • 2 inds; H3; 4-14.06.2014; FT; leg: IPT.

Genus: Paraclytus Bates, 1884

Paraclytus sexguttatus (Adams, 1817)

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT. 1 ind; H2; 5-15.05.2014; MT. 4 inds; H3; 5-15.05.2014; MT. 1 ind; H4; 5-15.05.2014; MT. 1 ind; H3; 15-25.05.2014; MT. 1 ind; H6; 25.06-5.07.2014; MT.

Genus: Parmena Dejean, 1821

Parmena pontocircassica Danilevsky et Miroshnikov, 1985*

GEORGIA • 1 ind; H3; 4-14.06.2014; MT. 1 ind; H3; 17-21.07.2019; HC; leg: JB.

Genus: Phymatodes Mulsant, 1839

Phymatodes femoralis Ménétriés, 1832*

GEORGIA • 5 inds; H3; 4-14.06.2014; FT; leg: IPT.

Genus: Prionus Geoffroy, 1762

Prionus coriarius (Linnaeus, 1758)*

GEORGIA • 1 ind; N41.85, E46.23; 2.07.2011; HC; leg: JH.

Genus: Rhagium Fabricius, 1775

Rhagium fasciculatum Faldermann, 1837*

GEORGIA • 1 ind; H1; 2-12.04.2014; MT. 3 inds; H1; 12-23.04.2014; MT. 1 ind; H2; 12-23.04.2014; MT. 1 ind; H1; 23.04-3.05.2014; MT. 3 inds; H2; 4.05.2014; MT. 5 inds; H2; 5-15.05.2014; MT. 2 inds; H3; 5-15.05.2014; MT. 2 inds; H4; 5-15.05.2014; MT.

Genus: Rhaesus Motschulsky, 1875

Rhaesus serricollis (Motschulsky, 1838)

GEORGIA • 1 ind; N41.84, E46.28; 30.04.2014; dead on beech snag (Fig. 1D); leg: JH. 1 ind; N41.84, E46.28; 1.05.2014; dead on beech snag; HC; leg: RP. 8 inds; N41.84, E46.28; 17-21.07.2019; HC; leg: JB, AB, JP.

Genus: Rosalia Audinet-Serville, 1833

Rosalia alpina (Linnaeus, 1758)

GEORGIA • 2 inds; H3; 14.07-5.08.2014; FT; leg: IPT. 1 ind; H3; 24.08.2011; HC; leg: GJ. 6 inds; H3; 17-21.07.2019; HC; leg: JB, AB, JP.

Genus: Rutpela Nakane & Ohbayashi, 1957

Rutpela maculata manca (Schaufuss, 1863)*

GEORGIA • 1 ind; H3; 15-25.06.2014; MT.

Genus: Stenocorus Geoffroy, 1762*

Stenocorus insitivus insitivus (Germar, 1824)

GEORGIA • 1 ind; N41.85, E46.29; 17-21.07.2019; HC; leg: AB.

Genus: Stenostola Dejean, 1835

Stenostola ferrea (Schrank, 1776)*

GEORGIA • 1 ind; H1; 12-23.04.2014; MT. 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H2; 4.05.2014; MT. 1 ind; H2; 5-15.05.2014; MT.

Genus: Stictoleptura Casey, 1924

Stictoleptura tonsa (J. Daniel et K. Daniel, 1891)*

GEORGIA • 1 ind; N41.85, E46.23; 2.07.2011; leg: JH.

Genus: Stromatium Audinet-Serville, 1834

Stromatium unicolor (Olivier, 1795)**

GEORGIA • 2 inds; 17-21.07.2019; HC; leg: JB.

Genus: Trichoferus Wollaston, 1854

Trichoferus campestris (Faldermann, 1835)**

GEORGIA • 1 ind; H3; 4-14.06.2014; FT. leg: IPT.

Genus: Xylotrechus Chevrolat, 1860

Xylotrechus arvicola (Olivier, 1795)*

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT.

X. rusticus (Linnaeus, 1758)

GEORGIA • 1 ind; N41.85, E46.23; 2.07.2011; HC; leg: JH.

Family: Cerylonidae*

Genus: Cerylon Latreille, 1802

Cerylon ferrugineum Stephens, 1830**

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 2 inds; H2; 1.05-14.06.2014; PT; leg: IPT.

Family: Ciidae*

Genus: Cis Latreille, 1797

Cis festivus (Panzer, 1793)

GEORGIA • 2 inds; H1; 23.04-3.05.2014; MT. leg: IPT.

C. fusciclavis Nyholm, (1953)**

GEORGIA • 2 inds; H2; 5-14.07.2014; FT; leg: IPT.

C. rugulosus Mellié, 1849**

GEORGIA • 3 inds; H1; 23.04-3.05.2014; MT. 10 inds; H1; 1.05-14.06.2014; HC; PT. 1 ind; H1; 15-25.06.2014; MT.

C. tomentosus Mellié, 1849*

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; N41.84, E46.28; 17-21.07.2019; HC; leg: AB.

Genus: Octotemnus Mellié, 1847

Octotemnus rugosopunctatus Drogvalenko, 2002**

GEORGIA • 1 ind; H2; 5-14.07.2014; FT; leg: IPT.

Family: Cleridae*

Genus Opilo Latreille, 1802

Opilo mollis (Linnaeus, 1758)

GEORGIA • 3 inds; H3; 4-14.06.2014; FT; leg: IPT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT.

Genus: Thanasimus Latreille, 1806

Thanasimus formicarius (Linnaeus, 1758)**

GEORGIA • 1 ind; H3; 5-15.05.2014; MT. 1 ind; H1; 1.05-14.06.2014; HC; PT. 1 ind; H3; 15-25.05.2014; MT.
The species is enlisted in Georgian Biodiversity Database (Tarkhnishvili and Chaladze 2022), whithout indication of voucher deposition or locality data.

Genus: Tillus Olivier, 1790

Tillus elongatus (Linnaeus, 1758) **

GEORGIA • 1 ind; H3; 15-25.06.2014; MT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT.

Family: Coccinelidae

Genus: Chilocorus Leach, 1815

Chilocorus renipustulatus (Scriba, 1791)*

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; N41.84, E46.28; 17-21.07.2019; leg: AB.

Family: Corylophidae*

Genus: Arthrolips Wollaston, 1854

Arthrolips picea (Comolli, 1837)

GEORGIA • 1 ind; H2; 12-23.04.2014; MT. 2 inds; H1; 23.04-3.05.2014; MT. 6 inds; H1; 1.05-14.06.2014; PT; leg: IPT. 2 inds; H1; 15-27.09.2014; MT.

Family: Cucujidae*

Genus: Laemophloeus Dejean, 1835

Laemophloeus monilis (Fabricius, 1787)**

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT.

Genus: Pediacus Shuckard, 1839

Pediacus dermestoides (Fabricius, 1793)**

GEORGIA • 1 ind; H1; 2-12.04.2014; MT. 2 inds; H1; 23.04-3.05.2014; MT.

Genus: Cucujus Müller, 1764

Cucujus haematodes caucasicus Motschulsky, 1845*

GEORGIA • 1 ind; H3; 5-15.05.2014; MT.

Family: Curculionidae

Genus: Anisandrus J.A. Ferrari, 1867

Anisandrus dispar (Fabricius, 1793)**

GEORGIA • 1 ind; H1; 2-12.04.2014; MT. 1 ind; H1; 3.05.2014; MT. 5 inds; H3; 5-15.05.2014; MT. 1 ind; H4; 5-15.05.2014; MT. 10 inds; H1; 15-25.05.2014; MT. 1 ind; H3; 14.06-5.07.2014; MT. 1 ind; H1; 15-25.08.2014; MT.

Genus: Hylesinus J.C. Fabricius, 1802

Hylesinus varius (Fabricius, 1775)**

GEORGIA • 1 ind; H1; 3.05.2014; MT.

Genus: Magdalis E.F. Germar, 1817

Magdalis fallax Kirsch, 1878*

GEORGIA • 1 ind; H2; 1.05-14.06.2014; PT; leg: IPT.

Genus: Phloeotribus P.A. Latreille, 1797

Phloeotribus scarabaeoides (Bernard, 1788)**

GEORGIA • 1 ind; H1; 23.04-3.05.2014; MT.

Genus: Platypus J.F.W. Herbst, 1793

Platypus cylindrus (Fabricius, 1793)**

GEORGIA • 9 inds; H2; 25.05-4.06.2014; MT. 8 inds; H2; 4-14.06.2014; MT. 1 ind; H3; 14.06-5.07.2014; FT. 4 inds; H1; 15-25.06.2014; MT. 7 inds; H3; 15-25.06.2014; MT. 1 ind; H1; 25.06-5.07.2014; MT. 3 inds; H1; 5-15.07.2014; MT. 1 ind; H1; 15-25.07.2014; MT. 1 ind; H1; 26.07-5.08. 2014; MT. 1 ind; H1; 15-25.08.2014; MT. 24 inds; H2; 15-25.08.2014; MT. 1 ind; H3; 4-14.09.2014; MT.

Genus: Scolytus E.L. Geoffroy, 1762

Scolytus intricatus Ratzeburg, 1837

GEORGIA • 1 ind; H3; 15-25.06.2014; MT.

Genus: Taphrorychus W.J. Eichhoff, 1878

Taphrorychus bicolor (Herbst, 1794)**

GEORGIA • 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT.

Genus: Trachodes E.F.Germar, 1823

Trachodes hystrix Gyllenhal, 1836*

GEORGIA • 1 ind; H1; 3.05.2014; MT. 1 ind; H1; 5-15.05.2014; MT. 1 ind; H3; 5-15.05.2014; MT. 1 ind; H3; 25.05-4.06.2014; MT. 1 ind; H3; 4-14.06.2014; MT. 1 ind; H1; 15-25.06.2014; MT. 1 ind; H1; 15-25.08.2014; MT.

Genus: Xyleborinus E.Reitter, 1913

Xyleborinus saxesenii Ratzeburg, 1837**

GEORGIA • 1 ind; H1; 5-15.05.2014; MT. 1 ind; H1; 15-25.05.2014; MT. 1 ind; H3; 14.06-5.07.2014; FT. leg: IPT.

Genus: Xylechinus F. Chapuis, 1869

Xylechinus pilosus (Ratzeburg, 1837)**

GEORGIA • 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H1; 5-15.07.2014; MT.

Genus: Xylosandrus E. Reitter, 1913

Xylosandrus germanus (Blandford, 1894)*

GEORGIA • 1 ind; H1; 5-15.05.2014; MT. 1 ind; H1; 15-25.05.2014; MT. 1 ind; H2; 25.05-4.06.2014; MT. 5 inds; H1; 5-15.07.2014; MT. 1 ind; H1; 5-15.08.2014; MT.

Family: Dasytidae

Genus: Dasytes Paykull, 1799

Dasytes niger (Linnaeus, 1760)**

GEORGIA • 1 ind; H5; 5-15.05.2014; MT.

Family: Elateridae

Genus: Ampedus Dejean, 1833

Ampedus ochropterus Germar, 1844*

GEORGIA • 1 ind; H3; 1.05-14.06.2014; FT. 3 inds; H3; 5-15.05.2014; MT.

A. pomorum (Herbst, 1784)*

GEORGIA • 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H2; 4.05.2014; MT. 3 inds; H3; 1.05-14.06.2014; FT. leg: IPT. 1 ind; H1; 5-15.05.2014; MT. 1 ind; H3; 5-15.05.2014; MT. 1 ind; H3; 15-25.05.2014; MT.

A. sinuatus Germar, 1844*

GEORGIA • 1 ind; H2; 1.05-14.06.2014; PT; leg: IPT.

A. wachtangi Dolin, 1970*

GEORGIA • 1 ind; H3; 15-25.06.2014; MT.

Genus: Anostirus C.G.Thomson, 1859

Anostirus castaneus (Linnaeus, 1758)**

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

Genus: Brachygonus du Buysson, 1912

Brachygonus cf. frater Wurst, 1995**

GEORGIA • 1 ind; H3; 5-15.05.2014; MT.

Genus: Cardiophorus Eschscholtz, 1829

Cardiophorus kryzhanovskyi Dolin & Chantladze, 1980

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT.

Genus: Denticollis Piller & Mitterpacher, 1783

Denticollis parallelicollis (Aubé, 1850)

GEORGIA • 1 ind; H2; 1.05-14.06.2014; PT; leg: IPT. 3 inds; H3; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H3; 25.05-4.06.2014; MT.

Genus: Hypoganus Kiesenwetter, 1858

Hypoganus stepanovi Denisova, 1948*

GEORGIA • 1 ind; H1; 25.05-4.06.2014; MT.

Genus: Lacon Laporte, 1838

Lacon punctatus (Herbst, 1779)

GEORGIA • 1 ind; N41.84, E46.28; 30.04.2014; at night on snag; leg: JH.

Genus: Melanotus Eschscholtz, 1829

Melanotus castanipes (Paykull, 1800)*

GEORGIA • 2 inds; H1; 3.05.2014; MT. 1 ind; H1; 5-15.05.2014; MT. 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

M. villosus (Geoffroy, 1785)*

GEORGIA • 1 ind; H3; 4.05-14.06.2014; MT. 1 ind; H3; 25.05-4.06.2014; MT.

Genus: Procraerus Reitter, 1905

Procraerus carinifrons (Desbrochers des Loges, 1875)*

GEORGIA • 1 ind; H3; 1.05-14.06.2014; FT; leg: IPT.

Family: Endomychidae*

Genus: Symbiotes Philip, 1956

Symbiotes gibberosus (Lucas, 1846)**

GEORGIA • 1 ind; H2; 4-14.06.2014; MT.

Family: Erotylidae

Genus: Dacne Latreille, 1797

Dacne semirufula (Reitter, 1897)*

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H1; 5-15.07.2014; MT. 4 inds; 17-21.07.2019; leg: JB, AB, JP.

Genus: Lycoperdina Latreille, 1807

Lycoperdina succincta (Linnaeus, 1767)**

GEORGIA • 1 ind; 17-21.07.2019; leg: AB.

Genus: Triplax Herbst, 1793*

Triplax aenea (Schaller, 1783)

GEORGIA • 1 ind; 17-21.07.2019; leg: AB.

Triplax elongata Lacordaire, 1842**

GEORGIA • 1 ind; 30.04.2014; at night on snag; HC; leg: JH.

Triplax rufipes (Fabricius, 1787)

GEORGIA • 1 ind; H1; 15-25.06.2014; MT. 11 inds; 17-21.07.2019; HC; leg: JB, AB, JP.

Genus: Tritoma Fabricius, 1775

Tritoma bipustulata Fabricius, 1775

GEORGIA • 1 ind; H1; 15-25.05.2014; MT.

Family: Eucnemidae*

Genus: Dromaeolus Kiesenwetter, 1858

Dromaeolus barnabita (Villa et Villa, 1838)**

GEORGIA • 1 ind; H2; 25.05-4.06.2014; MT.

Genus: Isorhipis Boisduval & Lacordaire, 1835

Isorhipis nigriceps (Mannerheim, 1823)**

GEORGIA • 14 inds; H1; 23.04-3.05.2014; MT. 4 inds; N41.84, E46.28; 30.04.2012; leg: RP. 6 inds; H3; 4-14.06.2014; FT; leg: IPT. 3 inds; H3; 1.05-14.06.2014; PT; leg: IPT. 3 inds; H2; 4.05.2014; MT. 12 inds; H1; 5-15.05.2014; MT. 6 inds; H2; 5-15.05.2014; MT. 5 inds; H3; 5-15.05.2014; MT. 3 inds; H1; 15-25.05.2014; MT. 1 ind; H3; 25.05-4.06.2014; MT. 1 ind; H3; 15-25.06.2014; MT.

Genus: Xylophilus Mannerheim, 1823

Xylophilus leseigneuri Olexa, 1960**

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

Family: Geotrupidae

Genus: Odonteus Samouelle, 1819*

O. armiger (Scopoli, 1772)**

GEORGIA • 2 inds; 15-25.05.2014; HC; leg: GJ. 1 ind; 25.05-4.06.2014; HC; leg: GJ.

Genus: Geotrupes Latreille, 1797

Geotrupes olgae (Olsoufieff, 1918)*

GEORGIA • 2 inds; 17-21.07.2019; HC; leg: JB, AB.

Family: Histeridae*

Genus: Onthophilus Leach, 1817

Onthophilus striatus (Forster, 1771)

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

Genus: Paromalus Erichson, 1834

Paromalus flavicornis (Herbst, 1791)

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

Family: Laemophlaeidae*

Genus: Laemophloeus Dejean, 1835

Laemophloeus monilis (Fabricius, 1787)

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT.

Family: Latridiidae*

Genus: Cartodere C.G.Thomson, 1859

Cartodere nodifer (Westwood, 1839)**

GEORGIA • 1 ind; H1; 15-25.08.2014; MT.

Genus: Corticaria Marsham, 1802

Corticaria alleni Johnson, 1974**

GEORGIA • 1 ind; H2; 5-14.07.2014; HC; FT.

Genus: Corticarina Reitter, 1880

Corticarina similata (Gyllenhal, 1827)*

GEORGIA • 2 inds; H3; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT. 2 inds; H2; 5-14.07.2014; FT; leg: IPT.

Genus: Enicmus C.G.Thomson, 1859

Enicmus brevicornis (Mannerheim, 1844)

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

Enicmus rugosus (Herbst, 1793)

GEORGIA • 2 inds; H1; 23.04-3.05.2014; MT. 2 inds; H1; 5-15.05.2014; MT. 13 inds; H1; 1.05-14.06.2014; PT; leg: IPT. 3 inds; H2; 1.05-14.06.2014; FT; leg: IPT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT. 24 inds; H2; 5-14.07.2014; FT; leg: IPT. 4 inds; H3; 14.07-5.08.2014; FT; leg: IPT.

Enicmus testaceus (Stephens, 1830)**

GEORGIA • 1 ind; H2; 1.05-14.06.2014; FT; leg: IPT. 1 ind; H2; 5-14.07.2014; FT; leg: IPT.

Genus: Latridius Herbst, 1793

Latridius consimilis (Mannerheim, 1844)

GEORGIA • 1 ind; H2; 5-14.07.2014; FT; leg: IPT.

L. hirtus Gyllenhal, 1827**

GEORGIA • 2 inds; H1; 1.05-14.06.2014; PT; leg: IPT. 2 inds; H2; 5-14.07.2014; FT; leg: IPT. 2 inds; H3; 14.07-5.08.2014; FT; leg: IPT.

L. minutus (Linnaus, 1767)

GEORGIA • 1 ind; H2; 5-14.07.2014; FT; leg: IPT.

L. porcatus (Herbst, 1793)**

GEORGIA • 2 inds; H2; 5-14.07.2014; FT; leg: IPT.

Genus: Melanophthalma Motschoulsky, 1866

Melanophthalma taurica (Mannerheim, 1844)**

GEORGIA • 3 inds; H1; 1.05-14.06.2014; PT; leg: IPT.

Genus: Stephostethus LeConte, 1878

Stephostethus alternans (Mannerheim, 1844)**

GEORGIA • 2 inds; H1; 23.04-3.05.2014; MT.

Stephostethus angusticollis (Gyllenhal, 1827)*

GEORGIA • 1 ind; H1; 5-15.05.2014; MT.

Family: Leiodidae*

Genus: Anisotoma Panzer, 1797

Anisotoma humeralis (Fabricius, 1793)**

GEORGIA • 1 ind; H1; 12-23.04.2014; MT. 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

Genus: Nemadus C.G. Thomson, 1867

Nemadus colonoides (Kraatz, 1851)**

GEORGIA • 1 ind; H4; 3.05.2014; MT. 1 ind; H3; 5-15.05.2014; MT. 1 ind; H1; 15-25.05.2014; MT. 3 inds; H3; 25.05-4.06.2014; MT. 1 ind; H3; 4-14.06.2014; MT. 2 inds; H1; 5-15.08.2014; MT. 2 inds; H1; 25.08-5.09.2014; MT. 1 ind; H1; 5-15.09.2014; MT. 2 inds; H1; 15-27.09.2014; MT.

Family: Lucanidae

Genus: Dorcus Macleay, 1819

Dorcus parallelipipedus (Linnaeus, 1758)

GEORGIA • 1 ind; H1; 26.07-5.08.2014; MT. 6 inds; 30.05.2012; HC; leg: AB. 6 inds; 17-21.07.2019; HC; leg: AB.

Genus: Lucanus Scopoli, 1763

Lucanus cervus (Linnaeus, 1758)

GEORGIA • 1 ind; H1; 25.06-5.07.2014; MT.

L. ibericus ibericus Motschulsky, 1845*

GEORGIA • ind; 24.08.2011; HC; leg: GJ. 17 inds; 17-21.07.2019; HC; leg: JB, AB, JP.

Genus: Platycerus Geoffroy, 1762

Platycerus perplexus Gusakov, 2003**

GEORGIA • 1 ind; H3; 2-12.04.2014; MT. 1 ind; H1; 12-23.04.2014; MT. 2 inds; H2; 12-23.04.2014; MT. 17 inds; H1; 3.05.2014; MT. 5 inds; H3; 9-15.05.2014; MT.

Genus: Sinodendron Hellwig, 1792

Sinodendron cylindricum (Linnaues, 1758)**

GEORGIA • 1 ind; H3; 15-25.06.2014; MT.

Family: Lymexylidae*

Genus: Hylecoetus Latreille, 1806

Elateroides dermestoides (Linnaeus, 1761)**

GEORGIA • 1 ind; H3; 15-25.05.2014; MT.

Family: Melandryidae*

Genus: Melandrya Fabricius, 1801

Melandrya caraboides (Linnaeus, 1760)

GEORGIA • 2 inds; H1; 23.04-3.05.2014; MT. 3 inds; H1; 1-14.05.2014; PT; leg: IPT. 5 inds; H1; 5-15.05.2014; MT. 1 ind; H3; 5-15.05.2014; MT. 2 inds; H3; 4-14.06.2014; FT; leg: IPT.

Melandrya rufipes Gebler, 1829**

GEORGIA • 1 ind; H1; 1-14.05.2014; PT; leg: IPT.

Genus: Orchesia Latreille, 1807

Orchesia undulata Kraatz, 1853

GEORGIA • 1 ind; H1; 15-25.05.2014; MT. 2 inds; H2; 25.05-4.06.2014; MT. 1 ind; H3; 4-14.06.2014; FT; leg: IPT.

Genus: Phloiotrya Stephens, 1832

Phloiotrya tenuis (Hampe, 1850)

GEORGIA • 3 inds; H3; 4-14.06.2014; FT; leg: IPT.

Family: Mordellidae*

Genus: Tomoxia Costa, 1853

Tomoxia bucephala Costa, 1854**

GEORGIA • 6 inds; H1; 1.05-14.06.2014; PT; leg: IPT. 2 inds; H1; 15-25.05.2014; MT. 2 inds; H1; 25.05-4.06.2014; MT. 1 ind; H2; 25.05-4.06.2014; MT. 1 ind; H1; 25.06-5.07.2014; MT.

Family: Mycetophagidae*

Genus: Eulagius Motschulsky, 1845

Eulagius acernus Motschulsky, 1845

GEORGIA • 2 inds; H1; 1.05-14.06.2014; PT; leg: IPT.

Genus: Litargus Erichson, 1846

Litargus connexus (Fourcroy, 1785)

GEORGIA • 1 ind; H3; 22-23.04.2014; MT. 6 inds; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H1; 5-15.05.2014; MT. 1 ind; H3; 5-15.05.2014; MT. 2 inds; H3; 25.05-4.06.2014; MT. 1 ind; H3; 14.06-5.07.2014; MT. 1 ind; H1; 5-15.08.2014; MT.

Genus: Mycetophagus Hellwig, 1792

Mycetophagus ciscaucasicus (Semenov, 1899)

GEORGIA • 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H3; 1.05-14.06.2014; PT; leg: IPT.

M. fulvicollis Fabricius, 1793

GEORGIA • 1 ind; H2; 12-23.04.2014; MT. 1 ind; N41.84, E46.28; 30.05.2012; at night on snag; leg: JH.

M. quadripustulatus (Linnaeus, 1760)*

GEORGIA • 1 ind; N41.84, E46.28, 30.05.2012; at night on fungi bodies, leg: TJ.

Family: Nitidulidae*

Genus: Cychramus Kugelann, 1794

Cychramus luteus (Fabricius, 1787)

GEORGIA • 5 inds; H2; 1.05-14.06.2014; PT; leg: IPT.

Genus: Cyllodes Erichson, 1843*

Cyllodes ater (Fabricius, 1787)

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; 17-21.07.2019, leg: AB, det. TG;

Genus: Epuraea Erichson, 1843

Epuraea aestiva (Linnaeus, 1758)

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H2; 1.05-14.06.2014; PT; leg: IPT.

E. melanocephala (Marsham, 1802)

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 5 inds; H2; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H3; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H1; 3.05.2014; MT. 1 ind; H1; 5-15.05.2014; MT. 1 ind; H3; 5-14.07.2014; FT; leg: IPT.

E. pallescens (Stephens, 1835)

GEORGIA • 2 inds; H2; 1.05-14.06.2014; PT; leg: IPT. 2 inds; H3; 1.05-14.06. 2014; PT; leg: IPT. 5 inds; H3; 5-14.07.2014; FT; leg: IPT.

Genus: Glischrochilus Reitter, 1873

Glischrochilus grandis (Tournier, 1872)

GEORGIA • 1 ind; H1; 3.05.2014; MT.

Genus: Ipidia Erichson, 1843

Ipidia sexguttata (Sahlberg, 1834)

GEORGIA • 1 ind; H3; 5-14.07.2014; FT; leg: IPT.

Genus: Soronia Erichson, 1843

Soronia grisea (Linnaeus, 1758)**

GEORGIA • 1 ind; H3; 5-14.07.2014; FT; leg: IPT.

Family: Oedemeridae*

Genus: Ischnomera Stephens, 1832

Ischnomera coerulea (Linnaeus, 1758)

GEORGIA • 2 inds; H3; 4-14.06.2014; FT; leg: IPT.

Family: Ptinidae*

Genus: Caenocara C.G.Thomson, 1859

Caenocara affine (Sturm, 1837)**

GEORGIA • 1 ind; H1; 5-15.07.2014; MT.

C. subglobosum (Mulsant & Rey, 1864)**

GEORGIA • 2 inds; H1; 5-15.07.2014; MT.

Genus: Hedobia Dejean, 1821

Hedobia pubescens (Olivier, 1790)**

GEORGIA • 1 ind; H1; 2-12.04.2014; MT. 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H3; 5-15.05.2014; MT. 1 ind; H3; 1.05-14.06.2014; FT; leg: IPT.

Genus: Hemicoelus LeConte, 1861

Hemicoelus fulvicornis (Sturm, 1837)**

GEORGIA • 1 ind; H3; 15-25.06.2014; MT.

Genus: Hyperisus Mulsant & Rey, 1863

Hyperisus plumbeum (Illiger, 1801)

GEORGIA • 1 ind; H3; 4.05.2014; MT.

Genus: Plumilus White, 1974

Plumilus grandicollis (Ménétriés, 1832)**

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H3; 14.06-5.07.2014; FTl leg: IPT.

Genus: Ptinomorphus Mulsant & Rey, 1868

Ptinomorphus rosti (Pic, 1896)**

GEORGIA • 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT. 1 ind; H3; 14.06-5.07.2014; FT; leg: IPT.

Genus: Ptinus Linnaeus, 1767

Ptinus biformis Reitter, 1880

GEORGIA • 1 ind; H3; 1.05-14.06.2014; FT; leg: IPT.

P. swaneticus Reitter, 1906

GEORGIA • 2 inds; H3; 15-25.06.2014; MT.

Genus: Xestobium Motschulsky, 1845

Xestobium rufovillosum (De Geer, 1774)**

GEORGIA • 1 ind; H3; 5-15.05.2014; MT.

Family: Pyrochroidae*

Genus: Pogonocerus Fischer von Waldheim, 1812

Pogonocerus thoracicus Fischer von Waldheim, 1812**

GEORGIA • 1 ind; H1; 15-25.06.2014; MT. 2 ind; H3; 15-25.06.2014; MT. 1 ind; H5; 15-25.06.2014; MT.

Genus: Pyrochroa Müller, 1764

Pyrochroa coccinea (Linnaeus, 1761)

GEORGIA • 1 ind; H1; 5-15.05.2014; MT. 2 inds; H2; 5-15.05.2014; MT.

Family: Rhysodidae**

Genus: Omoglymmius Ganglbauer, 1891

Omoglymmius germari (Ganglbauer, 1892)

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT.

Family: Salpingidae*

Genus: Salpingus Illiger, 1801

Salpingus caucasicus Reitter, 1905**

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H1; 3.05.2014; MT. 1 ind; H1; 15-27.09.2014; MT.

Family: Scarabaeidae

Genus: Acanthobodilus G. Dellacasa, 1983*

Acanthobodilus immundus (Creutzer, 1799)

GEORGIA • 1 ind; N41.84, E46.28; 17-21.07.2019; leg: AB.

Genus: Anomala Samouelle, 1819*

Anomala dubia abchasica Motschulsky, 1854

GEORGIA • 1 ind; N41.84, E46.28; 17-21.07.2019; leg: AB.

Genus: Cetonia Fabricius, 1775*

Cetonia aurata pallida (Drury, 1773)

GEORGIA • 10 inds; 24.08.2011; HC; leg: GJ. 1 ind; H3; 15-25.05.2014; MT. 7 inds; N41.84, E46.28; 17-21.07.2019; leg: AB.

Genus: Euoniticellus Janssens, 1953*

Euoniticellus fulvus (Goeze, 1777)

GEORGIA • 2 inds; N41.84, E46.28; 17-21.07.2019; leg: AB.

Genus: Maladera Mulsant & Rey, 1871

Maladera castanea (Arrow, 1913)**

GEORGIA • 3 inds; N41.84, E46.28; 17-21.07.2019; leg: AB.

Maladera punctatissima (Faldermann, 1835)

GEORGIA • 2 inds; H1; 15-25.05.2014; HC; leg: GJ. 2 inds; N41.84, E46.28; 17-21.07.2019; leg: AB.

Genus: Onthophagus Latreille, 1802*

Onthophagus coenobita (Herbst, 1783)

GEORGIA • 4 inds; N41.84, E46.27; 17-21.07.2019; leg: AB.

Onthophagus taurus (Schreber, 1759)

GEORGIA • 2 inds; N41.84, E46.27; 17-21.07.2019; leg: AB.

Genus: Oryctes Hellwig, 1798*

Oryctes nasicornis latipennis Motschulsky, 1845

GEORGIA • 3 inds; N41.84, E46.28; 17-21.07.2019; leg: JB, AB.

Genus: Polyphylla Harris, 1841*

Polyphylla fullo (Linnaeus, 1758)

GEORGIA • 2 inds; 24.08.2011; HC; leg: GJ.

Polyphylla olivieri (Laporte, 1840)

GEORGIA • 1 ind; 41.84, 46.27; 17-21.07.2019; leg: AB.

Genus: Protaetia Burmeister, 1842

Protaetia affinis pseudospeciosa (Medvedev, 1964)*

GEORGIA • 1 ind; 41.84, 46.27; 17-21.07.2019; leg: JB.

Protaetia speciosa speciosa (Adams, 1817)

GEORGIA • 1 ind; 41.84, 46.27; 17-21.07.2019; leg: AB.

Family: Silvanidae*

Genus: Uleiota Latreille, 1797

Uleiota planatus (Linnaeus, 1761)

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT. 1 ind; H1; 5-14.09.2014; MT.

Family: Sphindidae**

Genus: Aspidiphorus Dejean, 1821

Aspidiphorus lareyniei Jacquelin du Val, 1859

GEORGIA • 1 ind; H1; 5-15.07.2014; MT.

A. orbiculatus (Gyllenhal, 1808)

GEORGIA • 3 inds; H2; 4-14.06.2014; MT. 1 ind; H1; 15-25.07.2014; MT. 1 ind; H1; 25.08-5.09.2014; MT.

Family: Staphylinidae

Genus: Gyrophaena Mannerheim, 1831

Gyrophaena fasciata (Marsham, 1802)*

GEORGIA • 1 ind; H1; 15-25.05.2014; MT.

Genus: Lordithon C.G.Thomson, 1859

Lordithon trinotatus (Erichson, 1839)

GEORGIA • 1 ind; H3; 25.05-4.06.2014; MT.
Previously recorded by Japoshvili and Khachikov (2020).

Family: Tenebrionidae

Genus: Corticeus Piller & Mitterpacher, 1783

Corticeus unicolor Piller et Mitterpacher, 1783

GEORGIA • 1 ind; H1; 1.05-14.06.2014; PT; leg: IPT.

C. bicolor (Olivier, 1790)

GEORGIA • 1 ind; H3; 1.05-14.06.2014; FT; leg: IPT.

C. suberis (Lucas, 1864)*

GEORGIA • 2 ind; H3; 1.05-14.06.2014; FT; leg: IPT.

Genus: Cryphaeus Klug, 1833

Cryphaeus cornutus (Fisher von Waldheim, 1823)

GEORGIA • 1 ind; H3; 1.05-14.06.2014; FT; leg: IPT.

Genus: Diaclina Jacquelin du Val, 1861

Diaclina testudinea (Piller et Mitterpacher, 1783)

GEORGIA • 1 ind; N41.84, E46.28; 30.04.2014; at night on snag; leg: JH.

Genus: Diaperis Geoffroy, 1762

Diaperis boleti (Linnaeus, 1758)**

GEORGIA • 65 inds; 30.05.2012; at night on fungi (Fig. 1F); HC; leg: TJ.

Genus: Lagria Fabricius, 1775

Lagria hirta (Linnaeus, 1758)*

GEORGIA • 3 inds; H1; 15-25.06.2014; MT. 3 inds; H1; 5-14.09.2014; MT.

Genus: Metaclisa Jacquelin du Val, 1861

Metaclisa azurea (Waltl, 1838)

GEORGIA • 14 inds; N41.84, E46.28; 17-21.07.2019; leg: JB, AB, JP.

Genus: Neatus LeConte, 1862*

Neatus subaequalis Reitter, 1920**

GEORGIA • 8 inds; N41.84, E46.28; 17-21.07.2019; leg: JB, AB, JP.

Genus: Neomida Latreille, 1829

Neomida quadricornis (Motschulsky, 1873)

GEORGIA • 1 ind; N41.84, E46.28; 17-21.07.2019; leg: JB.

Genus: Platydema Laporte & Brullé, 1831

Platydema tristis Laporte & Brullé, 1831

GEORGIA • 6 inds; N41.84, E46.28; 17-21.07.2019; leg: JB, AB, JP.

Family: Throscidae**

Genus: Aulonothroscus Horn, 1890

Aulonothroscus brevicollis (Bonvouloir, 1859)

GEORGIA • 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H1; 15-25.05.2014; MT. 1 ind; H3; 4-14.09.2014; MT.

Genus: Trixagus Kugelann, 1794

Trixagus dermestoides (Linnaeus, 1767)

GEORGIA • 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H1; 5-15.05.2014; MT.

Family: Thymalidae**

Genus: Thymalus Latreille, 1802

Thymalus aubei Léveillé, 1877

GEORGIA • 1 ind; N41.84, E46.28; 17-21.07.2019; leg: AB.

Family: Trogossitidae*

Genus: Nemozoma Latreille, 1804

Nemozoma cornutum Sturm, 1826**

GEORGIA • 1 ind; N41.84, E46.28; 1.05.2014; leg: RP. 3 inds; H3; 1.05-14.06.2014; FT; leg: IPT.

Genus: Tenebroides Piller & Mitterpacher, 1783

Tenebroides mauritanicus (Linnaeus, 1758)**

GEORGIA • 1 ind; H3; 4-14.06.2014; FT; leg: IPT.
TThe species is enlisted in Georgian Biodiversity Database (Tarkhnishvili and Chaladze 2022), whithout indication of voucher deposition or locality data.

Family: Zopheridae*

Genus: Bitoma Herbst, 1793

Bitoma crenata (Fabricius, 1775)

GEORGIA • 1 ind; N41.84, E46.28; 30.04.2014; at night on snag; leg: JH; 1 ind; H1; 23.04-3.05.2014; MT. 1 ind; H1; 5-15.05.2014; MT.

Genus: Colydium Fabricius, 1793

Colydium filiforme Fabricius, 1793

GEORGIA • 1 ind; 30.04.2014; at night on snag; HC; leg: JH.

Genus: Endophloeus Dejean, 1834

Endophloeus exculptus Germar, 1847

GEORGIA • 8 inds; N41.84, E46.28; 30.04.2014; at night on snag; leg: JH. 4 inds; N41.84, E46.28; 30.04.2014; leg: RP.

Genus: Pycnomerus Erichson, 1842

Pycnomerus terebrans (Oliver, 1790)**

GEORGIA • 6 inds; N41.84, E46.28; 30.04.2014; at night on snag; leg: JH; 1 ind; N41.84, E46.28; 1.05.2014; leg: RP.

Genus: Synchita Hellwig, 1792

Synchita mediolanensis Villa et Villa, 1833**

GEORGIA • 1 ind; 30.04.2014; at night on snag; HC; leg: JH.

Synchita separanda (Reitter, 1882)**

GEORGIA • 2 inds; N41.84, E46.28; 30.04.2014; at night on snag; leg: JH.

Synchita undata (Guérin-Méneville, 1844)**

GEORGIA • 3 inds; H3; 1.05-14.06.2014; FT; leg: IPT.

Summary

During this survey, 195 species of coleopterans, belonging to 42 families and 172 genera, have been recorded from Lagodekhi Protected Areas. Thirty families are recorded for the first time from the studied area, three of which, Rhysodidae, Sphindidae, and Thymalidae, are new family records for Georgia. One hundred fifty-seven species are new to LPA and 66 of them are new country records. Given the high landscape and habitat diversity of LNP and Georgia, this study can be considered as a snapshot of coleopteran diversity even for LNP. Thus, it can be expected that further research will yield many new species for LNP and Georgia as a whole.

დასკვნა

ნაშრომში წარმოდგენილია ხეშეშფრთიანების კვლევის შედეგები ლაგოდეხის დაცული ტერიტორიებიდან. შეგროვებულ იქნა 42 ოჯახის 195 სახეობა, რომლებიც 172 გვარში არიან გადანაწილებული. 30 ოჯახი და 157 სახეობის ხოჭო საკვლევი ტერიტორიისათვის პირველადაა აღრიცხული, რომელთაგანაც 3 ოჯახი Rhysodidae, Sphindidae და Thymalidae და 66 სახეობა ახალია საქართველოს ფაუნისათვის.

Acknowledgements

The work of the first author was supported by the Shota Rustaveli National Science National Foundation under grant #FR/221/7-110/13. We thank the administration and staff of Lagodekhi protected areas and give special thanks to its director Mr Giorgi Sulamanidze, for his help and assistance during this project. We are grateful to the editor (Levan Mumladze) for extensive revision of the manuscript. Samples were collected under a permit (KA060264272737314, number 892) issued by the Georgian Agency of Protected Areas.

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